Catalogue of silviculture measures to improve forest biodiversity in protected areas, a summary

Jan Leugner, Karel Matějka

Biodiversity is a commonly used term, which is not exactly defined yet. It is based partly on another term being used for much longer time, diversity. For example, species diversity concept is an exactly measurable value within a biocoenosis (Magurran 2004). There are two components of the diversity (i) species richness and (ii) equitability. The former is a number of species within community while the latter expresses a share of particular species. To maintain biodiversity, species and/or whole communities are protected from extinction. From this point of view, red lists and red books comprising of many organisms are important. They are compiled to include either larger (for higher plants in the Czech Republic see Grulich 2012) or smaller regions (for South Bohemia see Chán et al. 1999, for the Šumava Mts. see Procházka et Štech 2002, for the Jeseníky Mts. see Bureš 2013).

Forests represent the principal climax ecosystem in the central Europe (Chytrý 2012). They are thus in the focus of nature conservation. However, there are is lack of primaeval forests nowadays. The protected forest areas have been changed and/or reshaped due to long-term human intervention, which is a thing the nature conservationists have to cope with.

The Catalogue of Silviculture Measures to improve Forest Biodiversity in Protected Areas (hereinafter referred as the Catalogue) adds more information to knowledge base of forest management in protected areas. Although relationships between both biodiversity and forest management have been studied a lot (Forkner et al. 2006, Gram et al. 2003), there is a need to propose and use so-called "non-standard" silvicultural measures. Central-European forests of lower (and middle) altitudes developed under several thousands of years lasting human interventions. The result is for instance light-demanding oak forests, which can be considered semi-natural ecosystems (see Chytrý 2012). This is also in accordance with archaeological findings such as distribution of prehistoric human settlements (Semotanová, Cajthaml et al. 2014) and potential vegetation map (Neuhäuslová et al. 1998). The forests were managed in such a way that relatively frequent removals of "fine" biomass contributed to existence of thin stands allowing penetration of solar radiation through broken canopy where spontaneous regeneration was common. At that time, wood was harvested using a selection cut (Dreslerová 2012). Contemporary Czech forestry abandoned the old management approaches. We have full canopy forest stands disabling long-term or permanent life of the species that benefitted from almost forgotten management systems. To maintain or restore the high level of biodiversity, the return to "the old-fashioned silviculture measures" is needed. Such measures aim to break the crown canopy (reduce stocking) which contributes to create conditions emulating ecotones i.e. transition zones between forested and non-forested areas. It is known that ecotones are typical of increased diversity. For example, birds use the opportunity to nest and forage there (Gram et al. 2003).

As for middle and upper forested sites, there were almost no human interventions in prehistoric period. The important human impact and following change of tree species share and forest structure date back to Medieval Age. Leaving forests to their spontaneous development (Kindlmann et al. 2012) and near-natural management emulating the forest gap dynamics are considered specific measures at these sites. Protected forest including NATURA 2000 areas can be managed specifically in order to achieve the objectives of protection. These specific approaches are usually a part of the management plans in the protected areas. The Catalogue describes eight measures that are potentially applicable for planning the care of forest ecosystems in order to preserve/improve their biodiversity. However, it is not a complete list of all possible measures. There are applicability of measures in terms of species, ecosystems and natural processes conservation including impact on forest production, methods to be used and feedback to the measure users to evaluate impact on biodiversity.

The Catalogue content is divided into three types of forest management:

  1. Non-intervention management using spontaneous development
    1. Leaving forest to spontaneous development
      Aim of measure is a protection of natural processes in protected forested areas being less-affected by human interventions in order to support and maintain natural dynamics and structure of ecosystems which is to keep site and forage conditions for species benefitting from the particular status of ecosystems.
  2. Near-natural management aiming to convert structure of managed forests close to the demanded structure being considered a pristine one
    1. Conversion into selection forest (or at least no clear-cutting system)
      Aim of measure is a conversion of clear-cutting into selection cutting system using a set of long-term applied management procedures. The measure focuses on protection of plants and animals bound to a growing-up developmental stage of temperate forest.
  3. Deliberate management supporting particular species or species communities being important from nature-conservation point of view
    1. Excessive thinning of forests
      Aim of measure is to maintain and improve biological diversity using forestry procedures creating and keeping a permanently broken canopy. Forestry follows needs of nature conservation though a certain economic benefits are possible. The measure is a part of "old-fashioned", mostly forgotten silvicultural systems such as coppicing, forest pasture etc.
    2. Coppice and coppice with standards systems
      Aim of measure is to create a manifold mosaic of forest stands within relatively small area with changing ecological conditions, which support many species with different demands. It focuses on light-demanding (intolerant) species particularly in lowlands and hilly areas. Simple coppicing can be combined with growing upper stratum of seed origin (standards).
    3. Forest pasture
      Aim of measure is to use an extensive pasture of domestic herbivores to increase and maintain biodiversity on forest and forest-steppe sites in protected areas.
    4. Overaged trees and wood debris management
      Aim of measure is leaving both lying and standing wood debris to be decomposed and inhabited spontaneously by saprophagous and the other organisms to improve forest biodiversity.
    5. Care of forest edges
      Aim of measure is to maintain a forest edges as typical ecotones in the landscape to stabilize adjacent forest stands and to improve biodiversity in cultural landscape. In addition, it supports both aesthetic and recreation values of landscapes.
    6. Intraspecies diversity care
      Aim of measure is to reveal potential problems concerning intraspecies diversity on example of important tree species on upper mountain sites, the Norway spruce (Picea abies). The measure focuses on both silvicultural and biological approaches to maintain and improve the intraspecies diversity.

The Catalogue is based on several working procedures. The first one included an analysis of protected areas care plans. The analysis showed what measures were proposed or applied in the protected forests. Next procedures' effect on biodiversity was studied from available literature. The impact of applied measures was evaluated in so-called demonstration sites i.e. research plots, which were established in different parts within the Czech Republic (Matějka et al. 2016).

 

Project Silviculture measures to improve forest biodiversity in protected areas
Project number EHP-CZ02-OV-1-015-2014, Supported by grant from Iceland, Liechtenstein and Norway